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Martinus Willem Woerdeman grad student 1931 Amsterdam (Neurotree)
 (Zur Entwicklung der Ganglienleiste)


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Pieter Dirk Nieuwkoop grad student 1946 Utrecht
Frits H. Sobels grad student 1952 Utrecht (FlyTree)
J. A. M. Van den Biggelaar grad student 1970 Utrecht (Evolution Tree)
Paulien Hogeweg grad student 1976 Utrecht (Computational Biology Tree)
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