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Franz von Leydig grad student 1877 Universität Bonn (Neurotree)
 (Die Nebenorgane des Auges der einheimischen Lacertidae)


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Jacob H.F. Kohlbrugge research assistant 1890 Amsterdam (Neurotree)
Eugene Dubois grad student 1897 (Neurotree)
Pieter Nicolaas van Kampen grad student 1904 Amsterdam
Lieven F. de Beaufort grad student 1908 Amsterdam (Systematics Tree)
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