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Charles Otis Whitman grad student 1894 Chicago
 (The embryology of the Unionidae.)


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Lewis Victor Heilbrunn grad student 1915 Chicago (Neurotree)
Edwin Joseph Cohn grad student 1917 Chicago (Chemistry Tree)
Carl Moore grad student 1919 Chicago (Neurotree)
Benjamin Harrison Willier grad student 1920 Chicago (Neurotree)
Emil Witschi post-doc 1926 Chicago
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