Keith Karoly, Ph.D.

Biology Reed College, Portland, OR 
plant mating systems, pollination biology, phylogeography, floral evolution
"Keith Karoly"


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Vanessa A. Koelling research assistant 2000-2002 Reed College
Brook T. Moyers research assistant 2006-2007 Reed College
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Conner JK, Issaka Salia O, Zhao ZG, et al. (2023) Rapid evolution of a family-diagnostic trait: artificial selection and correlated responses in wild radish, Raphanus raphanistrum. The New Phytologist
Waterman R, Sahli H, Koelling VA, et al. (2022) Strong evidence for positive and negative correlational selection revealed by recreating ancestral variation. Evolution; International Journal of Organic Evolution
Sapir Y, Karoly K, Koelling VA, et al. (2017) Effect of expanded variation in anther position on pollinator visitation to wild radish, Raphanus raphanistrum. Annals of Botany
Conner JK, Mills CJ, Koelling VA, et al. (2014) Artificial selection on anther exsertion in wild radish, Raphanus raphanistrum. Scientific Data. 1: 140027
Conner JK, Karoly K, Stewart C, et al. (2011) Rapid independent trait evolution despite a strong pleiotropic genetic correlation. The American Naturalist. 178: 429-41
Conner JK, Sahli HF, Karoly K. (2009) Tests of adaptation: functional studies of pollen removal and estimates of natural selection on anther position in wild radish. Annals of Botany. 103: 1547-56
Koelling VA, Karoly K. (2007) Self-pollen interference is absent in wild radish (Raphanus raphanistrum, Brassicaceae), a species with sporophytic self-incompatibility. American Journal of Botany. 94: 896-900
Matlaga D, Karoly K. (2004) Long-term grazing effects on genetic variation in Idaho fescue Journal of Range Management. 57: 275-279
McGlaughlin M, Karoly K, Kaye T. (2002) Genetic variation and its relationship to population size in reintroduced populations of pink sand verbena, Abronia umbellata subsp. breviflora (Nyctaginaceae) Conservation Genetics. 3: 411-420
Karoly K, Conner JK. (2000) Heritable variation in a family-diagnostic trait. Evolution; International Journal of Organic Evolution. 54: 1433-8
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