Thomas Richard Holtz, Ph.D.

Geology University of Maryland, College Park, College Park, MD 
Vertebrate Paleontology, Dinosaur Phylogenetics, Functional Anatomy, Paleobiogeography
"Thomas Holtz"


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Steven M Stanley research assistant 1983-1987 Johns Hopkins
 (Undergraduate advisor)
Leo J. Hickey grad student 1987-1992 Yale (Chemistry Tree)
 (Committee member)
John H. Ostrom grad student 1987-1992 Yale
 (Primary advisor)
Charles Lee Remington grad student 1987-1992 Yale
 (Member of committee)
Adolf Seilacher grad student 1987-1992 Yale University and University of Tuebingen
 (Graduate mentor, although not on committee)
Elisabeth S. Vrba grad student 1987-1992 Yale
 (On committee)


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Maria Eugenia Leone Gold research assistant University of Maryland
Susan Drymala research assistant 2005-2009 University of Maryland
Benjamin T Breeden research assistant 2008-2011 University of Maryland
Albert Chen research assistant 2012-2016 University of Maryland
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Snively E, O'Brien H, Henderson DM, et al. (2019) Lower rotational inertia and larger leg muscles indicate more rapid turns in tyrannosaurids than in other large theropods. Peerj. 7: e6432
Holtz TR. (2017) Nomenclature: Share names for dinosaur divisions. Nature. 545: 30
Brusatte SL, Carr TD, Williamson TE, et al. (2016) Dentary groove morphology does not distinguish 'Nanotyrannus' as a valid taxon of tyrannosauroid dinosaur. Comment on: "Distribution of the dentary groove of theropod dinosaurs: Implications for theropod phylogeny and the validity of the genus Nanotyrannus Bakker et al., 1988" Cretaceous Research
Holtz TR. (2014) Palaeontology: mystery of the horrible hands solved. Nature. 515: 203-5
Farlow JO, Holtz TR, Worthy TH, et al. (2013) Feet of the fierce (And not so Fierce): Pedal proportions in large theropods, other non-Avian Dinosaurs, and large ground birds Tyrannosaurid Paleobiology. 89-134
France CAM, Zelanko PM, Kaufman AJ, et al. (2007) Carbon and nitrogen isotopic analysis of Pleistocene mammals from the Saltville Quarry (Virginia, USA): Implications for trophic relationships Palaeogeography, Palaeoclimatology, Palaeoecology. 249: 271-282
Holtz TR. (2006) Palaeontology: a Jurassic tyrant is crowned. Nature. 439: 665-6
Holtz TR. (2005) All in the family Natural History. 114: 40-41
Holtz TR. (2004) Tyrannosauroidea The Dinosauria: Second Edition. 111-136
Holtz TR, Molnar RE, Currie PJ. (2004) Basal tetanurae The Dinosauria: Second Edition. 71-110
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