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Rudolf Leuckart grad student 1881-1881 Universität Leipzig
Karl Wilhelm von Kupffer grad student 1881-1882 Universität München (DevTree)
Karl Alfred von Zittel grad student 1882-1882 Universität München
 (Der Tarsus der Vögel und Dinosaurier)
Othniel Charles Marsh research scientist 1884-1890 Yale (Geotree)
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