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Otto Bütschli grad student 1902 Universität Heidelberg (DevTree)
 (Untersuchungen über Eireifung, Befruchtung und Zelltheilung bei Polystomum integerrimum Rud)
Richard Carl Wilhelm Theodor Ritter von Hertwig post-doc 1904 Universität München (Neurotree)
 (Habilitation: Der Chromidialapparat lebhaft funktionierender Gewebszellen)
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