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Rudolf Ludwig Karl Virchow research assistant 1859-1861 Universität Berlin (Neurotree)
Philipp Friedrich Arnold grad student 1860 Universität Heidelberg (Neurotree)
 (Die Bindehaut der Hornhaut und der Greisenbogen.)
Philipp Friedrich Arnold post-doc 1863 Universität Heidelberg (Neurotree)
 (Habilitation)
Nikolaus Friedreich research scientist 1863 Universität Heidelberg (Neurotree)
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