Parents

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Robert Almer Harper grad student 1904 UW Madison
 (Nuclear and cell division in the pollen-mother-cells of Lilium canadense.)
Eduard Adolf Strasburger post-doc 1904-1905 Universität Bonn (Marine Ecology Tree)

Children

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Gilbert Morgan Smith grad student 1913 UW Madison (Marine Ecology Tree)
Elizabeth McCoy grad student 1929 UW Madison (Microtree)
Delmer Clair Cooper grad student 1930 UW Madison
Sanford Soverhill Atwood grad student 1937 UW Madison
Robert W. Allard grad student 1946 UW Madison (Evolution Tree)