Parents

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Asa Gray post-doc 1872 Harvard

Children

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William T. Hornaday research assistant 1872-1873 Iowa State (Neurotree)
Cornelius Lott Shear research assistant 1897 University of Nebraska - Lincoln (Plant Biology Tree)
Fredric E. Clements grad student 1898 University of Nebraska
John Weaver grad student 1908 University of Nebraska
Melvin Randolph Gilmore grad student 1914 University of Nebraska (Plant Biology Tree)
Theodore Alexander Kiesselbach grad student 1918 University of Nebraska - Lincoln (Plant Biology Tree)
Joseph ​C Arthur grad student 1870-1973 Iowa State College (Evolution Tree)
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